Community-level network studies suggest that seed dispersal networks may share some universal properties with other complex systems. However, most of the datasets used so far in those studies have been strongly biased towards temperate birds, including not only dispersers, but also seed predators. Recent evidence from multi-taxon networks suggests that seed dispersal networks are not all alike and may be more complex than previously thought. Here, we used network theory to evaluate seed dispersal in a strongly impacted Atlantic Forest fragment in northeastern Brazil, where bats and birds are the only extant dispersers. We hypothesized that the seed dispersal network should be more modular then nested, and that the dispersers should segregate their services... |