The morphological and molecular evidence for higher-level reptile relationships is reassessed, A combined analysis of 176 osteological, 40 soft anatomical, and 2903 (1783 aligned) molecular characters in 28 amniote taxa yields the traditional reptile tree. Synapsids (including mammals) are the sister taxon to all other amniotes, including all extant reptiles. Turtles group with anapsid parareptiles and fall outside a monophyletic Diapsida. Within diapsids, squamates and Sphenodon form a monophyletic Lepidosauria, and crocodiles plus birds form a monophyletic Archosauria. This tree is identical to the tree strongly supported by the osteological data alone when fossils are included. In a combined analysis the strong osteological signal linking turtles with... |