The taxonomy of Hydractinia, Stylactaria and Podocoryna is discussed and the three genera are merged into Hydractinia since their diagnostic characters are liable to lead to polyphyly and paraphyly, due to repeated episodes of medusa reduction via heterochrony (paedomorphosis). The phylogeny of the Hydractiniidae is reconstructed by using two outgroups, Clava and Cytaeis, both having some characters in common with the Hydractiniidae. The resulting phylogenetic trees agree in identifying affinities among Hydractinia, Kinetocodium and Hydrocorella, all with polymorphic colonies with gastrozooids having oral tentacles. The position of Clavactinia (characterized by gastrozooids with widely scattered tentacles) is at the root of the tree if Clava is the... |