We explore the thesis that stochasticity in successional-disturbance systems can be an agent of species extinction. The analysis uses a simple model of patch dynamics for seral stages in an idealized landscape; each seral stage is assumed to support a specialist biota. The landscape as a whole is characterized by a mean patch birth rate, mean patch size, and mean lifetime for each patch type. Stochasticity takes three forms: (1) patch stochasticity is randomness in the birth times and sizes of individual patches, (2) landscape stochasticity is variation in the annual means of birth rate and size, and (3) turnover mode is whether a patch is eliminated by disturbance or by successional change. Analytical and numerical analyses of the model suggest that... |